In the distant future, humanity is forced to live in massive, mobile fortified cities due to fear of attack by giant monsters called Klaxosaurus. The Klaxosaurus can only be fought by Franks, giant robots that can only be piloted by a male and female pilot working together.
Despite being able to pilot the Streligia alongside Zero Two, Hiro has no memory of what happened in the cockpit, leading some of his fellow pilots to be skeptical about what really happened, as it is rumored that Zero Too is capable of piloting a Franks on his own. ,
After a mock battle, Hiro’s fellow pilots are assigned their first sortie with the task of defeating a Klaxosaur beneath the city, but just as they are about to leave, Ikuno and Mitsuru’s connections falter, thus Delphinum, Argent and Genista Let’s pilots proceed without them.
As Konrad squares Klaxosaurus hordes, forming a massive formation, near the plantation, the Parasites form a line of defense with Squad 26 at the front, Squad 13 in the middle, and Strelegia on the rear guard.
Due to their accomplishments in the previous battle, Zero Two is permanently assigned to Squad 13 and Hiro is officially recognized as a Franks pilot. Squad 13 are then rewarded with a holiday at a nearby beach. As Squad 13 plays, they observe Hiro’s relationship with Zero Two and begin to wonder what “kissing” and “love” really are.